The archaeocyathans occupied the nutrient
rich zone during the early Cambrian and there are no known occupants of
the nutrient poor zone. The rest of the Cambrian showed no significant
reef building activity during the rest of the Cambrian straight through
to the Early Ordivician. The Tabulate and Rugose corals originated
by calcification of some type of anemone like organism. The rugosoids
(as well as stromatopoids, spicular sponges and bryozoans) appeared
during the Silurian and Devonian in the form of great reef complexes.
Tabulate corals and stromatopoids occupied the nutrient poor zone during
this time. The rugosoids and tabulate corals were greatly affected
by the Late Devonian Extinction. During the Carboniferous the reef
niche was occupied until the Devonian. the niche was occupied by
calcified sponges and bryozoans at this time. The tabulates and rugosoids
hung on until late in the Paleozoic. Only to eventually succumb to
the forces of extinction at the end of the Permian. The Lower Triassic
rocks are void of fossil corals.
In the Middle triassic
the Scleractarians appeared from some unknown origin. By the Late
Triassic the Scleractarian had begun to contruct reef systems in the nutrient
poor zones. The Scleratarians were major reef builders for the rest
of Mesozoic and Cenozoic.
The evolutionary pattern
amongst the corals have not yet been established since the differences
in morphology have been a result of environmental alteration and
do not window into the genetic evolution of the corals. For this
reason micro evolutionary line lines of the corals are yet to be discovered.
There has been no undisputed claims as of yet.